Publications in peer-reviewed journals:


  • Oskyrko O., Mi C., Du W. (2024). Integrating the thermal dependence of sex ratio into distribution models to better predict the global invasion risk of the pond slider turtle, Trachemys scripta. Marine Life Science & Technology, (under review).
  • Oskyrko O., Mi C., Shai M., Du W. (2024). ReptTraits: a comprehensive dataset of ecological traits in reptiles. Scientific Data, 11, 243. Download PDF
  • Oskyrko O., Lysenko R., Obikhod D., Yakovliev M., Haidash O. (2024). Reptiles and their conservation in south-west Ukraine. Oryx, 2024: 1-10. Download PDF
  • Oskyrko O., Parry N., Jablonski D. (2024). Saint Patrick, what about legless lizards? Tracing the mitochondrial affiliation and possible origin of Anguis (Squamata: Anguidae) population in Ireland. Amphibia-Reptilia, 45(2024): 99-106.


  • Sreelatha L.B., Pérez i de Lanuza G., Oskyrko O., Gomes V., Andrade P., Boratyński Z., Carretero M.A. (2023). Towards a generalization of the behavioral trends in locomotion across color morphs of European wall lizard (Podarcis muralis). Acta Herpetologica, 18(2): 139-145. Download PDF
  • Mi C., Ma L., Yang M., Li X., Meiri S., Roll U., Oskyrko O., Pincheira-Donoso D., Harvey L., Jablonski D., Safaei-Mahroo B., Ghaffari H., Smid J., Jarvie S., Mwangi R., Masroor R., Kazemi S., Nneji L., Fokoua A., Taboue G.C.T., Bauer A., Nogueira C., Meirte D., Chapple D., Das I., Grismer L., Avila L., Ribeiro-Junio M., Oliver P., Torres-Carvajal O., Wagner P., Ron S., Wang Y., Itescu Y., Wilcove D., Nagy Z., Liu X., Du W. (2023). Global Protected Areas as refuges for amphibians and reptiles under climate change. Nature Communications, 14, 1389. Download PDF


  • Oskyrko O., Sreelatha L.B., Silva-Rocha I., Sos T., Vlad S.E., Cogălniceanu D., Stănescu F., Iskenderov T.M., Doronin I.V., Lisičić D., Carretero M.A. (2022). Molecular analysis of recently introduced populations of the Italian wall lizard (Podarcis siculus). Acta Herpetologica, 17(2): 147-157. Download PDF
  • Oskyrko O., Sreelatha L.B., Hanke G.F., Deichsel G., Carretero M.A. (2022). Origin of introduced Italian wall lizards, Podarcis siculus (Rafinesque-Schmaltz, 1810) (Squamata: Lacertidae), in North America. BioInvasions Records, 11(4): 1095–1106. Download PDF
  • Oskyrko O., Sos T., Vacheva E., Vlad S.E., Cogălniceanu D., Uller T., Feiner N., Carretero M.A. (2022). Unravelling the origin of the common wall lizards (Podarcis muralis) in south-eastern Europe using mitochondrial evidence. Biodiversity Data Journal, 10: e90337. Download PDF
  • Oskyrko O., Lysenko R., Jablonski D. (2022). The sheltopusik (Pseudopus apodus) in southwestern Ukraine? Insights from the museum collection. Evolutionary Systematics, 6(1): 71–76. Download PDF


  • Jablonski D., Sillero N., Oskyrko O., Bellati A., Čeirāns A., Cheylan M., Cogălniceanu D., Crnobrnja-Isailović J., Crochet P.A., Crottini A., Doronin I., Džukić G., Geniez P., Ilgaz Ç., Iosif R., Jandzik D., Jelić D., Litvinchuk S., Ljubisavljević K., Lymberakis P., Mikulíček P., Mizsei E., Moravec J., Najbar B., Maciej P., Pupins M., Sourrouille P., Strachinis I., Szabolcs M., Thanou E., Tzoras E., Vergilov V., Vörös J., Gvoždík V. (2021): The distribution and biogeography of slow worms (Anguis, Squamata) across the Western Palearctic, with an emphasis on secondary contact zones. Amphibia-Reptilia (published online ahead of print 2021). 
  • Čeirāns A., Gravele E., Gavarane I., Pupins M., Mezaraupe L., Rubenina I., Kvach Y., Skute A., Oskyrko O., Nekrasova O., Marushchak O., Kirjushina M. (2021). Helminth communities in amphibians from Latvia, with an emphasis on their connection to host ecology. Journal of Helminthology, 95: E48. Download PDF
  • Oskyrko O., Jablonski D. (2021): The first comprehensive data on the distribution of reptiles within the Southern Bug eco-corridor, Ukraine. Herpetozoa, 34:97–114. Download PDF


  • Oskyrko O. & Jablonski D. (2020): Swimming Behavior in a Caspian Whipsnake, Dolichophis caspius (Gmelin 1789). IRCF Reptiles & Amphibians 27: 277-278. Download PDF
  • Kuzmin Y., I. Dmytrieva, O. Marushchak, S. Morozov-Leonov, O. Oskyrko & O. Nekrasova (2020): Helminth species and infracommunities in frogs Pelophylax ridibundus and P. esculentus (Amphibia: Ranidae) in northern Ukraine. Acta Parasitologica, 65(2): 341-353. Download PDF
  • Oskyrko O., Laakkonen H., Silva-Rocha I., Jablonski D., Marushchak O., Uller T., Carretero M. (2020): The possible origin of the common wall lizards Podarcis muralis (Laurenti, 1768) in Ukraine. Herpetozoa, 33: 87-93. Download PDF
  • Udovychenko I., Oskyrko O., Marushchak O., Halenova T., Savchuk O. (2020): Identification of biologically active fractions in the dermal secretions of the genus Bombina (Amphibia: Anura: Bombinatoridae). Acta Herpetologica 15(1): 21-29. Download PDF


  • Marushchak O.Yu., Nekrasova O.D., Oskyrko O.S., Voitenko V.S., Zhytnyk D.O. (2019): Impact of Electromagnetic Field on Peculiarities of Rana temporaria Linnaeus, 1758 (Anura, Ranidae) Ontogeny. Journal of Environmental Research, Engineering and Management, 75 (2): 82-89. Download PDF

Other publications:

  • Garin-Barrio I., Blanco Y., Cabido C., Carretero M.A., Fernández-Arrieta A., Izagirre-Egaña A., López-Aizpuru J., Mandiola E., Oskyrko O., Silva-RochaI. (2020):Introducción reciente de Podarcis sicula en dos enclaves costeros del País Vasco. Bol. Asoc. Herpetol. Esp., 31(2): 181-186. Download PDF
  • Kyriachenko Y., Oskyrko O., Udovychenko I., Halenova T. (2020). Hemolytic activity of skin secretions of amphibians that inhabit the Ukraine territory. Bulletin of Taras Shevchenko National University of Kyiv-Biology, 80 (1): 6-10. Download PDF
  • Akulenko, N.M., Dziubenko, N.V., Marushchak, O.Yu., Nekrasova, O.D., Oskyrko, O.S.  (2019): Histological   Changes   in   Common   Toad, Bufo bufo (Anura, Bufonidae), Liver Tissue   under Conditions of Anthropogenically Transformed Ecosystems. Vestnik Zoologii, 53(6): 501–506. Download PDF
  • Udovychenko I., Dudkina Y., Oliinyk D., Oskyrko O., Marushchak O., Halenova T., Savchuk O. (2019): In vitro haemostatic effect of amphibian crude skin secretions in rabbit blood plasma. Journal of Biological Research, 92: 84-75. Download PDF
  • Nekrasova O.D., Oskyrko O.S., Marushchak O.Yu. (2018): Color features of sand lizards, Lacerta agilis (Sauria, Lacertidae), in Kyiv region (Ukraine). Vestnik Zoologii. 52(6): 495-500. Download PDF
  • Oskyrko O., Dudkina J., Nikolaieva I., Halenova T., Marushchak O. (2018): Variability and properties of host defense peptides from the skin secretions of Anurans. Bulletin of Taras Shevchenko National University of Kyiv. Biology: 2(76): 51-56. Download PDF
  • Nikolaieva I., Oliinyk D., Oskyrko O., Marushchak O., Halenova T., Savchuk O. (2018). Amphibian skin secretions: a potential source of proteolytic enzymes. Biotechnologia Acta 11(5): 42-48. Download PDF
  • Balashov I., Shvydka Z., Vasyliuk O., Marushchak O., Shyriaeva D., Oskyrko O. (2018): The first record of an invasive Caucasian land snail Harmozica ravergiensis in Central Ukraine (Stylommatophora, Hygromiidae). Ruthenica, 28(1): 43-45. Download PDF
  • Balashov I., Vasyliuk O., Shyriaieva D., Shvydka Z., Oskyrko O., Marushchak O., Stetsun H., Bezsmertna O., Babytskij A., Kostiushyn V. (2018): Terrestrial Molluscs in the Dry Grasslands of the Dnipro Upland (Central Ukraine): New Records, Rare Species and Conservation Potential. Vestnik Zoologii. 52(1): 3–12. Download PDF
  • Tytar V., Nekrasova O., Pupina A., Pupins M., Oskyrko O. (2018). Long-term bioclimatic modelling the distribution of the fire-bellied toad, Bombina bombina (Anura, Bombinatoridae), under the influence of global climate change. Vestnik Zoologii, 52 (4), 341-348. Download PDF